@article {Kohler2010,
	title = {Cryptic species and hybridization in the Anolis polylepis complex, with the description of a new species from the Osa Peninsula, Costa Rica (Squamata: Polychrotidae).},
	journal = {Zootaxa},
	volume = {2718},
	year = {2010},
	pages = {23 {\textendash} 38},
	abstract = {<p>Based on differences in hemipenial morphology we recognize two species of anoles related to Anolis polylepis: Anolis polylepis (Pacific versant of central and southern Costa Rica and western Panama, excluding the Osa Peninsula) and a species described herein which is restricted to the Osa Peninsula. The two species differ in hemipenial morphology (hemipenis bilobed in A. polylepis versus unilobed in the species from the Osa Peninsula) but show no discernable differences in external morphology (i.e., morphometrics, scalation, coloration, male dewlap). We therefore consider them to be cryptic species. At the neck of the Osa Peninsula where the ranges of the two species meet we detected a narrow (about 1 km wide) hybridization zone in which only individuals with an intermediate hemipenial morphology occur.</p>
},
	keywords = {Anolis, Anolis osa, Central America, cryptic species, hybridization, new species, Polychrotidae, Reptilia, Squamata},
	author = {K{\"o}hler, Gunther and Dehling, D Matthias and K{\"o}hler, Johannes}
}
@article {Kohler2010,
	title = {A revision of the Central American species related to Anolis pentaprion with the resurrection of A. beckeri and the description of a new species (Squamata: Polychrotidae).},
	journal = {Zootaxa},
	volume = {2354},
	year = {2010},
	pages = {1{\textendash}18},
	abstract = {<p>Based on differences in coloration, morphometrics, and scalation, I recognize seven species of anoles related to Anolis pentaprion in Central America: Anolis beckeri (southeastern Mexico to northern Nicaragua), A. cristifer (Pacific lowlands of Chiapas to central Guatemala), A. fungosus (Caribbean premontane zone from central Costa Rica to western Panama), A. pentaprion (Caribbean lowlands of southern Nicaragua to northwestern Colombia, also on Pacific versant in central and eastern Panama), A. salvini (Pacific premontane and lower montane zones from central Costa Rica to western Panama), A. utilensis (restricted to Isla de Utila, Honduras), and a new species (from the Pacific lowlands of Costa Rica and western Panama) described herein. The new species differs from A. pentaprion by having a red male dewlap with few large, widely spaced gorgetal scales scattered across the dewlap (pink with purple pigment between rows of gorgetals, and with regular rows of numerous small, narrowly spaced gorgetal scales in A. pentaprion). Furthermore, I resurrect the name A. beckeri Boulenger for the pentaprion-like populations of anoles distributed from southeastern Mexico to northern Nicaragua. Anolis beckeri differs from A. pentaprion most obviously in the dorsal tail scalation (a serrated caudal crest present in A. pentaprion versus no such crest in A. beckeri) and dewlap scalation (gorgetal rows with 17\&ndash;25 narrowly spaced scales in A. pentaprion versus gorgetal rows with 4\&ndash;10 widely spaced scales in A. beckeri). Finally, SMF 83608 is proposed as the neotype for Anolis pentaprion. I provide standardized descriptions of A. beckeri, A. pentaprion, and the new species described herein.</p>
},
	keywords = {Anolis, Anolis charlesmyersi, Central America, neotype designation, new species, Polychrotidae, Reptilia, Squamata},
	author = {K{\"o}hler, Gunther}
}
