E. E.  Williams, ?South American Anolis (Sauria, Iguanidae): two new species of the Punctatus group.?, Breviora, vol. 233, pp. 1?15, 1965.

G.  Köhler, Dehling, M. D., and Köhler, J., ?Cryptic species and hybridization in the Anolis polylepis complex, with the description of a new species from the Osa Peninsula, Costa Rica (Squamata: Polychrotidae).?, Zootaxa, vol. 2718, pp. 23 ? 38, 2010.

G.  Köhler, ?A revision of the Central American species related to Anolis pentaprion with the resurrection of A. beckeri and the description of a new species (Squamata: Polychrotidae).?, Zootaxa, vol. 2354, pp. 1?18, 2010.