G.  Köhler and Smith, E. N., ?A New Species of Anole of the Norops schiedei group from Western Guatemala (Squamata: Polychrotidae).?, Herpetologica, vol. 64, pp. 216?223, 2008.

G.  Köhler, Dehling, M. D., and Köhler, J., ?Cryptic species and hybridization in the Anolis polylepis complex, with the description of a new species from the Osa Peninsula, Costa Rica (Squamata: Polychrotidae).?, Zootaxa, vol. 2718, pp. 23 ? 38, 2010.

G.  Köhler, ?A revision of the Central American species related to Anolis pentaprion with the resurrection of A. beckeri and the description of a new species (Squamata: Polychrotidae).?, Zootaxa, vol. 2354, pp. 1?18, 2010.